Uncapped duo George Taylor and Charlie Shiel have been called up to the Scotland squad for the Six Nations clash against Wales along with D'Arcy Rae.

Gregor Townsend drafted in centre Taylor, scrum-half Shiel and prop Rae for the round-two encounter at Murrayfield on Saturday between two sides on a high following winning starts.

Rae earned his only cap against Ireland in 2019, coming off the bench in a 22-13 loss.

Ewan Ashman, Blair Kinghorn, Jamie Dobie and Rufus McLean have been released back to their clubs.

Scotland dominated defending champions England on the opening day of the tournament, ending a 38-year wait for a win at Twickenham with an 11-6 victory on Saturday.

Captain Stuart Hogg said they will remain grounded after regaining the Calcutta Cup.

The full-back, who was named man of the match in the defeat of Eddie Jones' side, said: "We are not going to get ahead of ourselves. We will enjoy this one, no doubt, but then it is on to Wales next week.

"We'll get back on the horse and ready to go because next week is another challenge for us and one we're really excited about."

Gregor Townsend praised the "calmness" and "togetherness" that saw Scotland end 38-years of Twickenham hurt, as Eddie Jones took responsibility for England's Calcutta Cup defeat.

Defending Six Nations champions England were caught cold by a dominant Scotland showing at an empty Twickenham, a venue where the visitors had not beaten their rivals since 1983.

Stuart Hogg and Finn Russell were influential in a game marking the 150th anniversary of this fixture, while rookie Cameron Redpath impressed and Duhan van der Merwe scored the game's only try in an 11-6 win.

Reflecting on a famous victory with ITV Sport, Scotland coach Townsend said: "I think there was just a calmness about how we played, a togetherness and an effort level that was superb. 

"We had a really good feeling with the way we trained and bonded. But you never know, it's the first game of the championship, there were a few who had their first game for a while or debuts. It was above expectations but what a start to the championship.

"It's impressive, to win here is very tough. Our line-out was superb, we really respect England's set-piece, their line-out has been outstanding the last couple of years.

"We put pressure on lines-outs, the scrums were good – then we were accurate in attack. It's difficult in these conditions to go wide, which we did. The kicking was excellent and so was the players' leadership – you could hear them in this empty stadium."

Captain Hogg revelled in the win but insisted the focus quickly turns to next week's home clash against Wales.

"It's amazing for us, I think we can't control what happened in the past that's gone, we can control what happens in the future," Hogg said, 

"We were a credit to a man, credit to England they provided a tough challenge. We stood up to it.

"We got excited during the week, we knew we could come down here and put in a performance. We looked after the ball in attack, went for multi-phase and caused England problems. We'll enjoy this one then it's onto Wales."

England also lost their 2020 Six Nations opener to France but rebounded to win a championship delayed by the coronavirus pandemic.

The hosts were left lacking in both the possession stats and penalty counts, and head coach Jones said the buck lies with him.

"Firstly congratulations to Scotland, they played very well, played with a lot intent and a lot of spirt, we just had one of those days, maybe I didn't prepare the team right, we were a long way from our best," Jones said.

"You just have days like that. It's my responsibility, every time a team goes on the pitch it's a head coach's responsibility."

England's midfield pairing of Henry Slade and rookie Ollie Lawrence struggled to get into the game but Jones had a simple reason to explain that.
"You've just got to look at the possession stats, they had 75 per cent possession, a penalty count of 10 to one. That's why the centres didn't touch the ball, it's the basics of rugby," he added.

"It is what it is, we get the players, we accept responsibility for the players, I accept responsibility for not getting players ready for this game."

Eddie Jones questioned whether Scotland could handle the "weight of expectation" and they provided the sweetest of answers by ending a 38-year wait for a win at Twickenham.

Time and again Scotland had failed to beat their fierce rivals in their own backyard, but that elusive victory finally came as they regained the Calcutta Cup on a wet Saturday evening in London.

Gregor Townsend's side dominated the Six Nations champions on the opening day of the tournament, winning 11-6 to leave England head coach Jones with a face like thunder.

Jones will be asking why his ill-disciplined side started the defence of their title with such a flat, insipid performance in a game that marked the 150th anniversary of rugby's oldest fixture

Scotland had produced a sensational fightback to draw 38-38 at the same venue two years ago, before being denied an astonishing victory late on.

They never looked like suffering more heartbreak on this occasion, Stuart Hogg leading by example as they won at the famous stadium for the first time since 1983 to leave England shellshocked.

Scotland certainly did not resemble a team who might be feeling the pressure as they bossed the game from start to finish.

The Red Rose, starting the tournament with a depleted pack, were guilty of indiscipline time and again, with referee Andrew Brace losing patience when he sent Billy Vunipola to the sin bin.

Finn Russell deservedly put Scotland in front with a penalty early on and almost set up a try for Duhan van der Merwe with a clever kick, but the leaping wing was unable to grab a high bouncing ball and touch down.

Van der Merwe was not to be denied soon after, fending off Mark Wilson's tackle to put Scotland 8-0 up on the half-hour mark, but Scotland suffered a blow when Russell was yellow-carded just before the break for tripping Ben Youngs.

The boot of Farrell reduced the deficit to two points at the interval, with Scotland surely heading to the dressing room thinking they should have been further ahead after being frustrated by resolute England defending.

Russell returned with Scotland still leading and they continued to boss possession, managing the game superbly, and the fly-half's second penalty put them 11-6 up before he missed another shot at goal.

A furious Jones marched from the stands to the touchline to try and turn the tide, replacing Jamie George and Youngs with Luke Cowan-Dickie and Dan Robson before the hour-mark.

The excellent Hogg kept them on the back foot with a sublime, mammoth kick into the corner - not for the first time - and England were warned over their penalty count again, but more desperate defence denied Scotland a second try as they continue to hammer at the door.

Lacklustre England's day was summed up when Jonny May knocked on under no pressure in the closing stages.

Hogg said Scotland felt ready to "create a little bit of history" and start a "new chapter" this weekend and, as they finally celebrated on the Twickenham turf, it was evident the Red Rose had failed to live up to expectations.

Scotland ended a 38-year wait for victory over England at Twickenham as a hard-working and industrious display earned a deserved 11-6 triumph and Calcutta Cup glory.

Not since 1983 had Scotland beaten their biggest rivals at the home of English rugby but the fired-up visitors outbattled and outplayed Eddie Jones' defending Six Nations champions.

Finn Russell and Stuart Hogg were the attacking catalysts in a match marking the 150th anniversary of the first meeting between these old foes.

Rookie Scotland centre Cameron Redpath showed why there is so much hype with a promising display in an empty stadium due to the coronavirus pandemic that delayed the finish to the 2020 championship.

England were without five forwards but even accounting for that the penalty count was much too high and they were lucky to only be 8-6 down at the break – Duhan van der Merwe crossing for the game's only try.

The end result is a third Calcutta Cup in four years for Scotland, who had previously lost nine in a row between 2008 and 2017.

England's ill-discipline allowed Russell to boot over an early penalty and an accumulation of infringements led to the sin-binning of Billy Vunipola for a high, albeit loose, arm on Scotland's number 10.

Buoyed by the extra man Hamish Watson was held up at the line by Jamie George and only a cruel bounce denied Van der Merwe from meeting Russell's expert kick.

Another bomb from Russell set in motion the move from which Van der Merwe bulldozed through Mark Wilson's tackle shortly after as Scotland had reward for their dominance.

But skipper Owen Farrell quickly kicked the hosts onto the board before Russell was binned for a crude trip on Ben Youngs and another three-pointer from England's captain brought the score within two.

The numerical disadvantage did little to curb Scotland's enthusiasm early in the second half and Russell kicked a routine penalty after Ellis Genge was harshly pinged for not attempting to bind with a low tackle.

Russell missed with another effort that he will feel he should have made but still England struggled to garner any momentum.

Stuart Hogg was also off target from the tee to make it a nervy ending, but the final whistle was met with shouts of joy from the visitors.

England's depleted pack lacks punch

There is a sense of deja vu for England who lost the opening match of their 2020 title-winning campaign to France.

The absence of Mako Vunipola and Kyle Sinckler clearly weakened a usually strong pack, while Joe Marler, Joe Launchbury and Sam Underhill were also absent. Without some key men, England lacked their usual set-piece prowess.

Russell and Hogg torment England

A year ago it looked as though Russell's international career may be over following a breakdown in the relationship with head coach Gregor Townsend.

His return to action for the 2020 finale was ended early due to an injury that ruled him out of the Autumn Nations Cup, but Scotland fans will be delighted their fly-half wizard has patched things up with Townsend after his magic helped seal a famous win.

He was helped by the brilliant Hogg, whose pin-point kicking and lightning-quick feet was a constant thorn in the side of an out-of-sorts England.

What's next?

England are at back at HQ as they aim to bounce back next Saturday when Italy visit Twickenham, the same day Scotland – who will now fancy a real crack at the title – host Wales at Murrayfield.

Scotland ended a 38-year wait for victory over England at Twickenham as a hard-working and industrious display earned a deserved 11-6 triumph and Calcutta Cup glory.

Not since 1983 had Scotland beaten their biggest rivals at the home of English rugby but the fired-up visitors outbattled and outplayed Eddie Jones' defending Six Nations champions.

Finn Russell and Stuart Hogg were the attacking catalysts in a match marking the 150th anniversary of the first meeting between these old foes.

Rookie Scotland centre Cameron Redpath showed why there is so much hype with a promising display in an empty stadium due to the coronavirus pandemic that delayed the finish to the 2020 championship.

England were without five forwards but even accounting for that the penalty count was much too high and they were lucky to only be 8-6 down at the break – Duhan van der Merwe crossing for the game's only try.

The end result is a third Calcutta Cup in four years for Scotland, who had previously lost nine in a row between 2008 and 2017.

England start their quest to retain the Six Nations title against Scotland on Saturday just over three months after they were crowned champions. 

The Red Rose dethroned Wales at the end of October, pipping France on points difference in a dramatic finale to a tournament that ended almost nine months after it started due to the COVID-19 pandemic. 

Eddie Jones' side are favourites to win the competition again, but Les Bleus are also well fancied to end an 11-year wait for Six Nations glory and face Italy in the first match. 

England do battle with Scotland for the Calcutta Cup at Twickenham, with Wales and Ireland locking horns in Cardiff on Sunday. 

Ahead of the opening round, we preview the upcoming matches with help from Opta.

ITALY v FRANCE

FORM

France have won 19 of their 21 previous Six Nations matches against Italy (L2), including their last seven in a row. Their two defeats in that time both came in Rome, in 2011 and 2013.  

Italy have lost their last 27 Six Nations matches, the longest losing streak in the history of the competition. Their last victory came at Murrayfield against Scotland in 2015, while they have not won at home since 2013. 

Les Bleus achieved a cumulative points difference of +31 in the first 20 minutes of their 2020 Six Nations matches. No other side managed a positive double-digit points difference in that period of matches, but Italy had a -39 difference. 

ONES TO WATCH

France wing Teddy Thomas scored a try in his last Test against Italy and is capable of lighting up the tournament. He can get off to a storming start in Rome.

Paolo Garbisi caught the eye at fly-half last year after being handed his debut against Ireland in October. Hopefully for his country, there is still much more to come from the 20-year-old.

ENGLAND v SCOTLAND

FORM

The last time England hosted Scotland in the Six Nations they drew 38-38 two years ago. Scotland clawed back the biggest ever half-time deficit (-24 pts, 7-31) by a side to avoid defeat in the tournament’s history, almost snatching victory before a late George Ford try denied them. 

Scotland have never beaten England at Twickenham in the Six Nations (D1, L9), their last victory against them the Red Rose at the London venue came in 1983 in the Five Nations (22-12).  

Gregor Townsend's side concluded the 2020 edition by winning their last three games in a row, their best run in the Championship since Italy joined in 2000. 

ONES TO WATCH

England captain Owen Farrell has scored exactly 1,000 Test points, one of just six men to reach that milestone. He is just four points away from reaching a century against Scotland.

Finn Russell returns to the Scotland side and the creativity provided by the fly-half will be key for Townsend's men.

WALES v IRELAND

FORM

Wales have lost their last two home games in the Six Nations, the last time they suffered three consecutive defeats at home was in 2002-03. Wayne Pivac's side have lost their last four games in the competition, their worst streak since 2006-07 (L5).  

No team scored more tries than Ireland in the Six Nations last year (17, along with France), four of those were launched from the back of a scrum. Wales (2) were the only other side to score multiple tries following a scrum.  

Ireland scored seven tries in the final quarter of their fixtures in 2020, more than any other side. Wales, meanwhile, scored the joint-second most in that period (6, level with England). 

ONES TO WATCH

Louis Rees-Zammit turned 20 this week and the Wales wing can celebrate in style with a livewire display at the Principality Stadium.

Ireland centre Garry Ringrose had terrible luck with injuries last year, but he is ready to fire on all cylinders in 2021.

Kyle Sinckler will miss England's opening game in this year's Six Nations after the prop was handed a two-week ban for swearing at a referee.

The Bristol Bears forward was cited by an independent commissioner for failing to respect the authority of a match official during Saturday's Premiership game against Exeter Chiefs.

Sinckler contested the charge at a disciplinary hearing, during which he accepted he had used foul language.

The 27-year-old was handed a suspension by an independent panel and will now be unavailable until February 9, three days after England open their 2021 campaign against Scotland.

"The player was candid in his evidence as to why he had done so and regretted his actions," a statement from the independent panel read.

"The panel found that his actions disrespected the authority of the referee and it was in breach of a core value of rugby - respect of match officials - and warranted a red card.

"The panel determined that in all the circumstances it was a low entry point with no relevant mitigation. The sanction is a two-week ban."

Following on from the Calcutta Cup clash at Twickenham, Eddie Jones' squad host Italy on February 13.

The Six Nations is scheduled to go ahead amid the ongoing coronavirus pandemic, though the European Champions Cup and Challenge Cup have been temporarily suspended after French authorities ruled the participation of Top 14 clubs to be too much of a public health risk.

French Rugby Federation (FFR) president Bernard Laporte says there is "no question" that the Six Nations will be staged this year.

The Telegraph this weekend reported that the tournament was in doubt after the French sports ministry imposed a ban on playing cross-border elite sports with clubs from the United Kingdom due to a new strain of coronavirus.

With the European Champions Cup and Challenge Cup reportedly set to be suspended, the French government and Six Nations officials are expected to hold talks on Monday.

Yet Laporte does not believe there is any doubt that the competition, which starts in just four weeks' time, can be staged. 

He told RMC on Sunday: "It's a puzzle, but I think we should not be alarmed for the Six Nations tournament.

"The tournament will be played, with a health protocol dictated by the government, and linked to this mutant virus. Everything went well in the autumn [when the Autumn Nations Cup was staged], so will the upcoming Six Nations tournament, I'm no more worried than that."

The former Les Bleus coach added: "We have a meeting at the beginning of the week with the ministry of sports, a meeting also with the Six Nations Council.

"But we already talked to each other every day, and today there is no question of considering anything, except that we will play the tournament."

France are due to face Italy in the first game of the Six Nations at Stadio Olimpico on February 6.