Scotland ended a 38-year wait for victory over England at Twickenham as a hard-working and industrious display earned a deserved 11-6 triumph and Calcutta Cup glory.
Not since 1983 had Scotland beaten their biggest rivals at the home of English rugby but the fired-up visitors outbattled and outplayed Eddie Jones' defending Six Nations champions.
Finn Russell and Stuart Hogg were the attacking catalysts in a match marking the 150th anniversary of the first meeting between these old foes.
Rookie Scotland centre Cameron Redpath showed why there is so much hype with a promising display in an empty stadium due to the coronavirus pandemic that delayed the finish to the 2020 championship.
England were without five forwards but even accounting for that the penalty count was much too high and they were lucky to only be 8-6 down at the break – Duhan van der Merwe crossing for the game's only try.
The end result is a third Calcutta Cup in four years for Scotland, who had previously lost nine in a row between 2008 and 2017.
England's ill-discipline allowed Russell to boot over an early penalty and an accumulation of infringements led to the sin-binning of Billy Vunipola for a high, albeit loose, arm on Scotland's number 10.
Buoyed by the extra man Hamish Watson was held up at the line by Jamie George and only a cruel bounce denied Van der Merwe from meeting Russell's expert kick.
Another bomb from Russell set in motion the move from which Van der Merwe bulldozed through Mark Wilson's tackle shortly after as Scotland had reward for their dominance.
But skipper Owen Farrell quickly kicked the hosts onto the board before Russell was binned for a crude trip on Ben Youngs and another three-pointer from England's captain brought the score within two.
The numerical disadvantage did little to curb Scotland's enthusiasm early in the second half and Russell kicked a routine penalty after Ellis Genge was harshly pinged for not attempting to bind with a low tackle.
Russell missed with another effort that he will feel he should have made but still England struggled to garner any momentum.
Stuart Hogg was also off target from the tee to make it a nervy ending, but the final whistle was met with shouts of joy from the visitors.
England's depleted pack lacks punch
There is a sense of deja vu for England who lost the opening match of their 2020 title-winning campaign to France.
The absence of Mako Vunipola and Kyle Sinckler clearly weakened a usually strong pack, while Joe Marler, Joe Launchbury and Sam Underhill were also absent. Without some key men, England lacked their usual set-piece prowess.
Russell and Hogg torment England
A year ago it looked as though Russell's international career may be over following a breakdown in the relationship with head coach Gregor Townsend.
His return to action for the 2020 finale was ended early due to an injury that ruled him out of the Autumn Nations Cup, but Scotland fans will be delighted their fly-half wizard has patched things up with Townsend after his magic helped seal a famous win.
He was helped by the brilliant Hogg, whose pin-point kicking and lightning-quick feet was a constant thorn in the side of an out-of-sorts England.
What's next?
England are at back at HQ as they aim to bounce back next Saturday when Italy visit Twickenham, the same day Scotland – who will now fancy a real crack at the title – host Wales at Murrayfield.
'World-class' Scotland will provide Wallabies' sternest test – Skelton
Australia will face their sternest test to date when they continue their autumn tour against a "world-class" Scotland team at Murrayfield on Sunday.
That is the view of forward Will Skelton, who believes the Wallabies' victories in England and Wales have brought a new energy and confidence to their camp.
Having crashed out in the pool stage at last year's World Cup in France before finishing last in the Rugby Championship, Australia have looked a team reborn in recent weeks.
A first victory at Twickenham since 2015 and a 52-20 rout of Wales – their biggest-ever away win in matches between the teams – have made it an excellent tour to date for Joe Schmidt's side.
But La Rochelle lock Skelton feels their toughest assignment is to come this week, saying: "We've won two games, we're very aware of that.
"The Test against Scotland is going to be the hardest one yet. We're excited about this weekend and we're looking forward to ripping into it.
"Physically they're up there with the top five in the world. I think set-piece wise they're really strong.
"You've got a guy like Finn Russell who's a maestro at the back there. I think it's going be a close one and we'll be up for it.
"There's some class players in the Scottish team and, when they're at it they're world-class, so we're going to have to be on top of our game."
Wales must 'show courage and front up' against South Africa, says Gatland
Warren Gatland has urged Wales to "show real courage and front up" in their final Autumn Nations clash against South Africa.
Gatland's side bring the curtain down on a disappointing year when they welcome the reigning world champions to the Principality Stadium on Saturday.
Wales have lost an unprecedented 11 straight Tests, and could go an entire calendar year without tasting success for the first time since 1937.
Gatland has made four changes from last weekend's 52-20 defeat to Australia, with Sam Costelow, Rio Dyer, Christ Tshiunza and Taine Plumtree all coming into the starting XV.
Gareth Anscombe, Aaron Wainwright and Cameron Winnett drop out of the squad, while Adam Beard misses out through injury.
And Gatland wants his players to demonstrate fighting spirit against the Springboks.
"Last week's result hurts, and we are just as disappointed by it as the fans," he told reporters during his pre-match press conference.
"Our focus now is on training and preparing well for our final game of this Autumn Nations Series.
"There were good elements that we can definitely build on going into Saturday, but we have to improve our accuracy.
"We know what a quality side South Africa are and the physicality they bring. This week, we need to show real courage and front up against the world champions."
'Playing for England is the Mecca' according to Itoje amid global breakaway talk
Maro Itoje has reaffirmed his commitment to England amid talks of a proposed global breakaway league.
Steve Borthwick’s side are looking to end their five-match losing run when they face Japan in their final match of the autumn series on Sunday.
But there is fear that Borthwick could lose many of his players to the new league that reportedly are offering 40 £1m-a-season contracts for marquee players and more periods of rests for the players.
The Rugby Football Union (RFU) only select players for the national team from English clubs, meaning any player joining the breakaway league would jeopardise their Test career.
But Itoje has insisted that playing for England is the pinnacle for him and his teammates.
"I don't really know any of the details of it as yet, but wearing this rose is extremely important to me and my teammates," said Itoje.
"Playing for England is the Mecca."
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