Props Kyle Sinckler and Mako Vunipola have returned to the England squad for the Six Nations encounter with Italy at Twickenham on Saturday.
British and Irish Lions duo Sinckler and Vunipola missed the 11-6 Calcutta Cup defeat to Scotland on the opening day of the tournament due to suspension and an Achilles injury respectively.
They will be available to beef up the defending champions' pack against an Azzurri side smarting from a 50-10 hammering at the hands of a rampant France at Stadio Olimpico.
Harry Williams and Tom West will not feature for Eddie Jones' side after they were released to return to their clubs.
The Red Rose never got going in a poor performance versus dominant Scotland, but they won the tournament last year after losing their opening match to France and number eight Billy Vunipola said they can respond again.
He said: "It is frustrating. I won't lie. But this happened last year, and we managed to pull it back so it's massive for all of us to make sure we get around each other and whatever we need to fix this week we do as soon as possible.
"We only have a week to do it, which is probably the best thing for us. We don't have a fallow week to let that settle in and frustrate us.
"You never really forget these days, you never really forget these moments, and to be honest I don't really want to forget them.
"I want to move on from it, but I don't want to forget it as it keeps you sharp. Losing like this at Twickenham is never what you dream of."
'World-class' Scotland will provide Wallabies' sternest test – Skelton
Australia will face their sternest test to date when they continue their autumn tour against a "world-class" Scotland team at Murrayfield on Sunday.
That is the view of forward Will Skelton, who believes the Wallabies' victories in England and Wales have brought a new energy and confidence to their camp.
Having crashed out in the pool stage at last year's World Cup in France before finishing last in the Rugby Championship, Australia have looked a team reborn in recent weeks.
A first victory at Twickenham since 2015 and a 52-20 rout of Wales – their biggest-ever away win in matches between the teams – have made it an excellent tour to date for Joe Schmidt's side.
But La Rochelle lock Skelton feels their toughest assignment is to come this week, saying: "We've won two games, we're very aware of that.
"The Test against Scotland is going to be the hardest one yet. We're excited about this weekend and we're looking forward to ripping into it.
"Physically they're up there with the top five in the world. I think set-piece wise they're really strong.
"You've got a guy like Finn Russell who's a maestro at the back there. I think it's going be a close one and we'll be up for it.
"There's some class players in the Scottish team and, when they're at it they're world-class, so we're going to have to be on top of our game."
Wales must 'show courage and front up' against South Africa, says Gatland
Warren Gatland has urged Wales to "show real courage and front up" in their final Autumn Nations clash against South Africa.
Gatland's side bring the curtain down on a disappointing year when they welcome the reigning world champions to the Principality Stadium on Saturday.
Wales have lost an unprecedented 11 straight Tests, and could go an entire calendar year without tasting success for the first time since 1937.
Gatland has made four changes from last weekend's 52-20 defeat to Australia, with Sam Costelow, Rio Dyer, Christ Tshiunza and Taine Plumtree all coming into the starting XV.
Gareth Anscombe, Aaron Wainwright and Cameron Winnett drop out of the squad, while Adam Beard misses out through injury.
And Gatland wants his players to demonstrate fighting spirit against the Springboks.
"Last week's result hurts, and we are just as disappointed by it as the fans," he told reporters during his pre-match press conference.
"Our focus now is on training and preparing well for our final game of this Autumn Nations Series.
"There were good elements that we can definitely build on going into Saturday, but we have to improve our accuracy.
"We know what a quality side South Africa are and the physicality they bring. This week, we need to show real courage and front up against the world champions."
'Playing for England is the Mecca' according to Itoje amid global breakaway talk
Maro Itoje has reaffirmed his commitment to England amid talks of a proposed global breakaway league.
Steve Borthwick’s side are looking to end their five-match losing run when they face Japan in their final match of the autumn series on Sunday.
But there is fear that Borthwick could lose many of his players to the new league that reportedly are offering 40 £1m-a-season contracts for marquee players and more periods of rests for the players.
The Rugby Football Union (RFU) only select players for the national team from English clubs, meaning any player joining the breakaway league would jeopardise their Test career.
But Itoje has insisted that playing for England is the pinnacle for him and his teammates.
"I don't really know any of the details of it as yet, but wearing this rose is extremely important to me and my teammates," said Itoje.
"Playing for England is the Mecca."
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