Wales have been warned they do not hold a monopoly on passion as Will Stuart called on England to prove that fighting spirit is also part of their genetic make-up.
The rivals collide at Twickenham on Saturday with Steve Borthwick’s new-look team aiming to build on their winning start to the Guinness Six Nations against Italy in Rome.
While it is assumed England’s opponents will always play with emotional intensity, spurred on by facing the tournament’s most unpopular team, scrum coach Tom Harrison bristles at the idea that the passion flows in one direction only.
“It’s an interesting assumption to presume Wales would have an extra desire than us,” Harrison said.
“There is rivalry both ways. This England is a special group. They’ve trained and gelled really well together.”
England are presenting their first appearance at Twickenham since beginning their post-2023 World Cup rebuild as the start of a new era.
Apart from seeking to build on the greater enterprise shown in attack against Italy and continue to bed in their new defensive system, they want to forge a reputation as a side that will not take a backwards step.
“We beat Wales with 12 men here in August and that’s partly what we want our DNA to be – never stop fighting and whatever the circumstances, to come out on top,” Stuart said.
“Loads is made of the rivalry with Wales but they’re a great side and keep battling. That’s part of the DNA we pride ourselves on as well.
“We’re massively excited to put a positive stamp on Twickenham and cement it as a hard place to come and play.
“When the atmosphere is electric here it’s the best place I’ve ever played in. It’s on us as players to get the crowd going by playing in a dominant way.
“It’s on us to provide performances to push the crowd to spur us on. It’s always a big occasion against Wales, but we have to focus on ourselves.”
Captain Jamie George has urged his players to be unafraid to show their passion as England look to reconnect with supporters after years of underachievement in the Six Nations.
For Stuart, the Bath tighthead prop, facing Wales at Twickenham will be a moment of intense pride, even if that emotion is visible only in smaller moments.
“I don’t know if I am the most outwardly shouty or passionate bloke, but inside there’s a lot there,” he said.
“You might occasionally get a shout after a scrum penalty or something like that. But apart from that it’s catching my breath and going again!
“I like to be as relaxed as possible until we start strapping at the hotel and then it’s headphones on and be with myself a little bit.
“Some people are pretty relaxed until kick-off. It’s different for everyone.”
Steward and Van Poortvleit among England changes for Springboks Test
Freddie Steward and Jack van Poortvliet will return for England on Saturday, with Steve Borthwick making four changes against South Africa.
England were unchanged for their narrow defeats against New Zealand and Australia, but the Leicester pair, who were not included in the squad for either of those matches, come straight in to replace George Furbank and Ben Spencer.
Ollie Sleightholme scored his first international tries in the defeat to Australia and is rewarded with a maiden Test start in place of Immanuel Feyi-Waboso, while Sam Underhill comes in for Tom Curry, with both players missing due to head injuries.
England have lost each of their last four Test matches, their longest such run since losing five straight games in 2018.
"We're excited to challenge ourselves against the world's top-ranked team and back-to-back Rugby World Cup champions," Borthwick said.
"Test matches against South Africa are always thrilling contests, and I'm sure Saturday will be no exception."
Meanwhile, South Africa have made 12 changes to their starting XV from their 32-15 win over Scotland on Sunday.
Captain Siya Kolisi and Pieter-Steph du Toit return in the back row, while Cheslin Kobe is also named in the side.
The world champions have got a more balanced look to their bench this time around though, with coach Rassie Erasmus opting not to name seven forwards among the replacements this time around.
England squad in full:
Freddie Steward, Tommy Freeman, Ollie Lawrence, Henry Slade, Ollie Sleightholme, Marcus Smith, Jack van Poortvliet, Ellis Genge, Jamie George (captain), Will Stuart, Maro Itoje, George Martin, Chandler Cunningham-South, Sam Underhill, Ben Earl.
Replacements: Luke Cowan-Dickie, Fin Baxter, Dan Cole, Nick Isiekwe, Alex Dombrandt, Harry Randall, George Ford, Tom Roebuck.
South Africa squad in full:
Alphelele Fassi, Cheslin Kolbe, Jesse Kriel, Damian De Allende, Kurt-Lee Arendse, Maine Libbok, Grant Williams, Ox Nche, Bongi Mbonambi, Wilco Loux, Eben Etzebeth, RG Snyman, Siya Kolisi (captain), Pieter-Steph Du Toit, Jasper Wiese.
Replacements: Malcolm Marx, Gerhard Steenekamp, Vincent Koch, Elrigh Louw, Kwagga Smith, Cobus Reinach, Handre Pollard, Lukhanyo Am.
Healy in line to equal Ireland cap record versus Argentina
Cian Healy is set to equal Brian O'Driscoll's Ireland cap record when they face New Zealand in Friday's autumn international in Dublin.
The 37-year-old prop has represented his country on 132 occasions and, after being named on the bench, will equal O'Driscoll's mark if introduced versus the Pumas.
Ireland suffered their first home defeat since 2021 last time out, going down by a 23-13 scoreline against New Zealand.
Despite that loss, head coach Andy Farrell has only made one change to Ireland's starting lineup, bringing Robbie Henshaw in at inside centre in place of Bundee Aki.
Argentina also named their team on Wednesday, with vice-captain Pablo Matera, Guido Petti and Matias Moroni replacing Santiago Grondona, Franco Molina and Matias Orlando.
Ireland team: Hugo Keenan; Mack Hansen, Garry Ringrose, Robbie Henshaw, James Lowe; Jack Crowley, Jamison Gibson-Park; Andrew Porter, Ronan Kelleher, Finlay Bealham; Joe McCarthy, James Ryan; Tadhg Beirne, Josh van der Flier, Caelan Doris.
Replacements: Rob Herring, Cian Healy, Thomas Clarkson, Ryan Baird, Peter O'Mahony, Craig Casey, Sam Prendergast, Jamie Osborne.
Argentina team: Juan Cruz Mallia; Rodrigo Isgro, Lucio Cinti, Matias Moroni, Bautista Delguy; Tomas Albornoz, Gonzalo Bertranou; Thomas Gallo, Julian Montoya, Joel Sclavi; Guido Petti, Pedro Rubiolo; Pablo Matera, Juan Martin Gonzalez, Joaquin Oviedo.
Replacements: Ignacio Ruiz, Ignacio Calles, Francisco Gomez Kodela, Franco Molina, Santiago Grondona, Gonzalo Garcia, Santiago Carreras, Justo Piccardo.
Pressure to win for England is a 'privilege', says Wigglesworth
Senior England coach Richard Wigglesworth believes that the pressure to win matches is part of the privilege of playing for the national team.
England face world champions South Africa at the Allianz Stadium on Saturday with displeasure growing after four straight defeats.
Included in that run were narrow losses in their first two Autumn Nations Series games against New Zealand and Australia.
Though England have won three of their last four games against South Africa at Allianz Stadium, Wigglesworth insisted everyone within the squad wants to improve.
"This job of being part of the England team is such a privilege, because so many people care, which makes your feeling on the matter really big," Wigglesworth told BBC Sport.
"Our job is to step back from that and see what was right, what was wrong, and what can we fix.
"We need to see the game for what it is, make the decisions for what they are, and get better every week.
"It is our job to make sure the players understand as much as they can that they can’t get affected by that as well.
"We know the attention the team gets is result-dependent, and rightly so."
The last time England and South Africa faced one another was in the Rugby World Cup semi-final in 2023, in which the Springboks came back from 15-6 down to win 16-15.
England will have to be wise to how the visitors have evolved since that game according to Wigglesworth.
"They have a squad and a plan they have done for a while, and when you are successful you tend to stick to things," he said.
"I have really enjoyed watching what they do, and how they do it. It is our job to meet that and rise to it."
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