Sam Simmonds and Jack van Poortvliet will start for England against Japan at Twickenham on Sunday.

Eddie Jones has named five changes from the shock 30-29 defeat to Argentina last weekend as they prepare to face Japan, who he coached between 2012 and 2015.

Number eight Simmonds has not featured for England since the Six Nations in March but comes in for Billy Vunipola, who drops to the bench.

Alex Coles is replaced by debutant David Ribbans at lock while Van Poortvliet, who scored within 30 seconds of coming on the pitch against Argentina, will start at scrum-half.

Jonny May also comes in for Joe Cokanasiga and Guy Porter is the final introduction as he is chosen to start over Manu Tuilagi at outside centre.

Owen Farrell will captain England and earn his 99th international cap, while hooker Jamie George is on the bench after recovering from a foot injury.

England team to face Japan: 15. Freddie Steward, 14. Jack Nowell, 13. Guy Porter, 12. Owen Farrell (c), 11. Jonny May, 10. Marcus Smith, 9. Jack van Poortvliet; 1. Ellis Genge, 2. Luke Cowan-Dickie, 3. Kyle Sinckler, 4. David Ribbans, 5. Jonny Hill, 6. Maro Itoje, 7. Tom Curry, 8. Sam Simmonds.

Replacements: 16. Jamie George, 17. Mako Vunipola, 18. Joe Heyes, 19. Alex Coles, 20. Billy Vunipola, 21. Ben Youngs, 22. Henry Slade, 23. Manu Tuilagi.

Danny Care has been left out of England's training squad ahead of the Autumn Nations Series but Manu Tuilagi and Sam Simmonds return.

Scrum-half Care featured in the Red Rose's series victory over Australia in July, but is not among the 36 players who will report for a three-day camp this weekend.

Centre Tuilagi and number eight Simmonds are back in the fold after recovering from injuries, while pivot Ben Youngs has been selected ahead of the November encounters with Argentina, Japan, New Zealand and South Africa.

Simmonds gets the nod amid reports that he could leave Exeter Chiefs to join Montpellier, a move that would make him ineligible to play for his country.

Locks Alex Coles and Hugh Tizard have been called up for the first time, while Alex Mitchell, Tom Pearson and David Ribbans are also included.

Courtney Lawes and Maro Itoje are among the absentees due to injury.

England head coach Eddie Jones said: "With a year to go to the Rugby World Cup, this is a big opportunity for players to come in and impress. We want them to show real energy and enthusiasm and that they want to be a part of this massive year.

"It doesn't mean that those who have been left out won't be considered for the Autumn Nations Series matches. We'll be looking at club games, form and fitness and the door is left open for those players.

"We finished the Australia tour well. It was a fantastic experience, particularly for the younger players. We now have to start again, but we'll build on what we've done there and continue that momentum."

England training squad:

Forwards: Ollie Chessum, Alex Coles, Luke Cowan-Dickie, Tom Curry, Ellis Genge, Jamie George, Joe Heyes, Jonny Hill, Lewis Ludlam, Tom Pearson, David Ribbans, Bevan Rodd, Patrick Schickerling, Sam Simmonds, Will Stuart, Hugh Tizard, Billy Vunipola, Mako Vunipola, Jack Walker, Jack Willis.

Backs: Henry Arundell, Joe Cokanasiga, Fraser Dingwall, Owen Farrell, Tommy Freeman, George Furbank, Will Joseph, Jonny May, Alex Mitchell, Jack Nowelll, Guy Porter, Marcus Smith, Freddie Steward, Manu Tuilagi, Jack van Poortvliet, Ben Youngs.

Alun Wyn Jones has been named captain of the British and Irish Lions for their tour of South Africa, leading a 37-man squad.

Warren Gatland announced his selection on Thursday ahead of the trip in July, which follows a warm-up against Japan next month.

Wales great Jones has been chosen for the leadership role having featured in the past nine Lions Tests going back to 2009, the longest run in the professional era.

Jones has already skippered the side once previously in the absence of the injured Sam Warburton in Australia eight years ago, clinching a first series win since 1997.

Elsewhere in Gatland's squad, Ireland's former World Rugby Player of the Year Johnny Sexton misses out despite playing a key role on the 2017 trip to New Zealand.

Billy Vunipola was also a notable absentee from the list, while other big calls saw Wales centre Jonathan Davies, England prop Kyle Sinckler and Ireland second row James Ryan excluded.

However, Gatland did find room for Exeter Chiefs number eight Sam Simmonds, who has not played a Test for England since March 2018.

"We believe we've picked a squad capable of winning a Test series in South Africa," Gatland - who opted for 11 Englishmen, 10 Welshmen and eight apiece from Ireland and Scotland - said following the squad announcement.

"Selecting a Lions squad is never easy and, in many ways, this has been the most challenging selection I have been involved in.

"Over the course of the last three weeks the coaches and I have rigorously debated each position.

"We saw some outstanding performances in the recent Six Nations, so competition for places has been tough with some incredibly tight calls to make. 

"However, we are very happy with the squad we have assembled and look forward to meeting up in Jersey in just over a month's time to start our preparation to take on the world champions.

"We've left out some very talented players which gives an indication to the strength of this squad and we know how important that stand-by list will be.

"Being selected for a Lions tour is the greatest honour for a British and Irish player and I congratulate everyone named today."

The Lions will face the Springboks in Tests on July 24, July 31 and August 7.

British and Irish Lions squad in full:

Josh Adams (Cardiff, Wales), Bundee Aki (Connacht, Ireland), Dan Biggar (Northampton Saints, Wales), Elliot Daly (Saracens, England), Gareth Davies (Scarlets, Wales), Owen Farrell (Saracens, England), Chris Harris (Gloucester, Scotland), Robbie Henshaw (Leinster, Ireland), Stuart Hogg (Exeter Chiefs, Scotland), Conor Murray (Munster, Ireland), Ali Price (Glasgow Warriors, Scotland), Louis Rees-Zammit (Gloucester, Wales), Finn Russell (Racing 92, Scotland), Duhan van der Merwe (Edinburgh, Scotland), Anthony Watson (Bath, England), Liam Williams (Scarlets, Wales); Tadhg Beirne (Munster, Ireland), Jack Conan (Leinster, Ireland), Luke Cowan-Dickie (Exeter Chiefs, England), Tom Curry (Sale Sharks, England), Zander Fagerson (Glasgow Warriors, Scotland), Taulupe Faletau (Bath, Wales), Tadhg Furlong (Leinster, Ireland), Jamie George (Saracens, England), Iain Henderson (Ulster, Ireland), Jonny Hill (Exeter Chiefs, England), Maro Itoje (Saracens, England), Alun Wyn Jones (Ospreys, Wales), Wyn Jones (Scarlets, Wales), Courtney Lawes (Northampton Saints, England), Ken Owens (Scarlets, Wales), Andrew Porter (Leinster, Ireland), Sam Simmonds (Exeter Chiefs, England), Rory Sutherland (Edinburgh, Scotland), Justin Tipuric (Ospreys, Wales), Mako Vunipola (Saracens, England), Hamish Watson (Edinburgh, Scotland).

Alun Wyn Jones has been named captain of the British and Irish Lions for their tour of South Africa, leading a 36-man squad.

Warren Gatland announced his selection on Thursday ahead of the trip in July, which follows a warm-up against Japan next month.

Wales great Jones has been chosen as captain having featured in the past nine Lions Tests going back to 2009, the longest run in the professional era.

Jones has already skippered the side once in the absence of the injured Sam Warburton in Australia eight years ago, clinching a first series win since 1997.