Glasgow back-rower Ally Miller has been given his first Scotland call-up ahead of Saturday’s Guinness Six Nations showdown with Ireland.

The 27-year-old, who initially struggled to establish himself with the Warriors after arriving at Scotstoun from Edinburgh in 2021, has been rewarded by Gregor Townsend for his excellent form under Franco Smith this term.

Miller joins up with the squad as fellow back-rower Hamish Watson drops out along with prop Jamie Bhatti and second-rower Glen Young. None of the forward trio were involved on Saturday as the Scots crashed to a damaging 31-29 defeat away to Italy.

An update released by Scottish Rugby on Monday afternoon also revealed that 21-year-old Glasgow second-rowers Alex Samuel and Max Williamson are training with the squad.

Their more experienced Warriors second-row colleague Scott Cummings, meanwhile, is adamant that Scotland will have no Rome hangover as they prepare for their championship-concluding match away to Ireland.

Had the Scots won in Italy at the weekend, they would be heading to Dublin this weekend for a title shootout with their hosts.

Instead, the visit to the Irish capital has become an exercise in saving face for Townsend’s under-pressure team.

If Scotland – currently third in the table – lose on Saturday, they will end the tournament with just two wins from five and could feasibly finish fifth if Italy defeat Wales this weekend.

“It’s going to be a tough game,” said Cummings. “To get a win out there we’re going to have to have a 10 out of 10 performance.

“Ireland are one of the best teams in the world and we’ve got to go to Dublin, and try and put a performance in.

“We need to change one or two things (from Italy) and make sure we finish this campaign the way we want to and represent the country how we want to with the final performance.”

Scotland have not won in Dublin since 2010 and have lost each of their last nine meetings with Ireland, including a chastening 36-14 World Cup pool-stage defeat in Paris just five months ago.

Asked how easily it would be for the Scots to rouse themselves for such a formidable match just a week after one of their worst results of Townsend’s near-seven-year reign, Cummings said: “Quite easily, to be honest. In the big games the mentality takes care of itself.

“We remember that last game at the World Cup when we didn’t show up how we wanted to show up and that still lives in our memories.

“There’s going to be no shortage of motivation after Saturday’s game and the last time we played Ireland so we’re excited for it.

“It’s obviously going to be a tough challenge but one we’re ready to take on.”

Huw Jones batted away any notion that Scotland head coach Gregor Townsend was under pressure after Saturday’s Guinness Six Nations implosion in Italy.

The Scots blew their chance to set up a shootout for the title with Ireland in Dublin next weekend as they went down 31-29 in Rome despite holding a 22-10 lead after an encouraging opening half an hour.

It was the Azzurri’s first Six Nations victory at home for 11 years and their first in the championship since winning in Wales two years ago.

The surprise defeat cranks up the heat on Townsend just five months after the Scots – widely deemed to have one of the best squads in their history at present – suffered a second consecutive World Cup group-stage exit on the 50-year-old’s watch.

However, Jones claimed the players should carry the can for the debacle in the Eternal City and appeared irritated by the suggestion that it would place the long-serving head coach under renewed scrutiny.

“I don’t know about that,” said the experienced centre. “We’re all behind the coaches, we’re all behind Gregor.

“We love the way we play, the way we want to play. We have a good plan.

“When we execute it, it’s brilliant and we play some good rugby. I don’t think this defeat was on Gregor, I think it was on the players.

“We didn’t execute our plan well enough and Italy played well.”

Scotland looked in control after three tries in the opening half hour from Zander Fagerson, Kyle Steyn and Pierre Schoeman. But a disallowed George Horne touchdown – after a foul in the build-up by Schoeman was detected – two minutes into the second half when they led 22-16 proved pivotal.

Italy, who had scored in the first half through Martin Page-Relo, turned the screw with tries from debutant Louis Lynagh and substitute Stephen Varney, and some excellent kicking under pressure from Paolo Garbisi took the game away from the Scots before Sam Skinner’s late try gave them a glimmer of what ultimately proved false hope.

Saturday was one of Jones’ most soul-destroying days in a Scotland jersey, and he said: “We hate losing. It’s really tough to take, hugely disappointing.

“We spoke during the week before the game about having our best performance, having an 80-minute performance, but we were really poor in the second half. We let the game slip away from us.

“Credit to Italy, they were good, but we had that try chalked off and then conceded four or five penalties in a row. We couldn’t get back in the game and they managed that period better than us.

“Our discipline wasn’t good enough. We didn’t react to that try-swing well enough.

“We are gutted with our performance. Across the board we managed it badly. The leadership and the processes and the communication was good but we’ve all got to look at ourselves and the actions we took.”

While the manner of the defeat itself was bad enough, there was further reason for Scottish frustration later in the day when Ireland’s surprise defeat to England meant Townsend’s men had effectively squandered a golden chance to set up a last-day title shootout with Ireland.

Had they won in Rome, they would have been able to secure a first championship triumph since 1999 with victory in Dublin next weekend.

Instead – although still with an unrealistic mathematical chance of the title – they head to the Irish capital scrambling to avoid a demoralising two-win, bottom-half finish from a campaign that previously promised so much.

Jones admitted it felt like Scotland had let a huge opportunity slip from their grasp.

“Yes, definitely,” he said. “It is really disappointing.

“We wanted to get a win and then go to Dublin next week full of confidence and try to do something but this obviously takes the wind out of the sails a bit.

“We’ve got to react, we’ve got to react quickly and prepare for another game.

“We’ll go through a range of emotions but we’ve got to review it objectively and then turn our attention to Ireland.”

Frustrated Gregor Townsend refused to entertain questions about his future after Scotland suffered a calamitous second-half collapse on their way to a costly 31-29 Guinness Six Nations defeat in Italy.

The Scots arrived in Rome aiming to enhance their bid for a first top-two finish in the championship this century, while they still held out some hope of winning the title. However, their loss in the Eternal City, allied to England’s win over Ireland, leaves them staring at the prospect of a bottom-half finish.

Things looked to be going well for Townsend’s team when they led 14-3 and 22-10 during the first half following tries by props Zander Fagerson and Pierre Schoeman and wing Kyle Steyn. A disallowed George Horne try early in the second half following a foul by Schoeman proved pivotal though.

Italy, who had scored in the first half through Martin Page-Relo, turned the match in their favour with tries from debutant Louis Lynagh and substitute Stephen Varney. And some excellent kicking under pressure from Paolo Garbisi took the game away from the Scots before Sam Skinner’s late try gave them a glimmer of hope.

Scotland incredibly still have an outside chance of finishing in the top two as they prepare to head to Ireland for their final match next weekend, but losing to Italy for the first time since 2015 represents a major setback for Townsend just five months after an underwhelming display at the World Cup brought pool-stage elimination.

When the head coach, who is contracted until 2026, was asked if he felt his job was under threat after his side became the first to lose a Six Nations match in Rome since 2013, he said: “I’m not going to answer that question.

“We’re disappointed with the result today, but we know this team have come on since the World Cup.

“We can look at this result and be really disappointed, but I’ve been really proud of how this team’s played during the championship so far.

“I believe in this group. You can look at results and say: ‘We’ve lost to Italy, this team aren’t going to take that next step’. Or you can look at where we’ve grown since the World Cup.

“I still believe the performances the players are putting in can show we can beat anybody.

“But if we take our eye off the ball for a period of time, we can be beaten. That’s what happened today.”

Townsend admitted Scotland were architects of their own downfall – although he did acknowledge a superb performance from Italy, who won their first championship match since their victory in Wales two years ago.

“The credit has to go to Italy as well,” said Townsend. “There are two teams that play the game, not just one. I thought that Italy were very good even in the first half where we dominated most of the possession.

“But our actions after the try that was disallowed weren’t good enough. We gave Italy a way into the game through penalties and field position. If you give that to any team in the Six Nations, they’re going to take the points, which they did.

“It’s very frustrating that we didn’t get that try, which is down to our execution, and after that it was even more frustrating.”

Townsend admitted the result “hurts a lot” but he remains steadfast in his belief that Scotland are continuing to make progress overall under his stewardship.

“We won in Wales for the first time since 2002 this year,” he pointed out. “We retained the Calcutta Cup. We had a decision (over what would have been a match-winning try) go against us in the game with France.

“This result is very disappointing. We’ve lost against a good Italy side, one of the best Italian teams I’ve seen. But today is not what defines this team.”

Scotland’s Guinness Six Nations campaign unravelled in disastrous fashion in Rome as they crashed to a first defeat against Italy since 2015.

Gregor Townsend’s side looked set to tighten their grip on second place – and keep themselves on course for a first top-two finish this century – when they held 14-3 and 22-10 leads in the first half following tries from props Zander Fagerson and Pierre Schoeman either side of a score from Kyle Steyn.

However, the Scots completely lost their way after the interval as they fell to a 31-29 loss against an Italian side who – despite several encouraging displays that helped draw a capacity crowd to the Stadio Olimpico – had not won a Six Nations match since victory in Wales two years ago.

This chastening defeat in the Eternal City is sure to crank up the heat on head coach Townsend five months after exiting the World Cup at the group-stage.

Italy got the chance to get the scoreboard ticking over just seconds into the match when Scotland were penalised on their own 22 immediately from the kick-off. Paolo Garbisi saw the ball fall off the tee while the clock was running but the fly-half kept his composure to re-tee and send his kick between the posts.

Scotland soon gained a foothold, however, and they got themselves in front in the sixth minute when Fagerson powered his way over from close range after a sustained spell of pressure inside the Italian 22. Finn Russell converted.

The visitors crossed the whitewash again five minutes later when they worked the ball out to the right and Blair Kinghorn fed Steyn, who bundled his way past two Italians to cross the line. Russell converted again.

The Azzurri summoned a swift response as scrum-half Martin Page-Relo lobbed a clever kick over the top and centre Juan Ignacio Brex ran gleefully over just to the left of the posts in the 15th minute. Garbisi converted.

Shortly after Russell kicked a close-range penalty, Scotland scored their third try of the match in the 28th minute as Schoeman finished things off after being fed by George Turner in a driving maul. Russell’s conversion drifted wide.

Italy finished the first half strongly and reduced their interval deficit to 22-16 with a couple of penalties kicked by Garbisi and then Page-Relo, who was on target from close to the half-way line.

Scotland thought they had extended their advantage two minutes into the second period when scrum-half George Horne – on his first start since the 2019 World Cup – bolted over after excellent play by Huw Jones to release him but it was subsequently chalked off after Schoeman was adjudged to have committed a foul in the build-up.

The prop’s needless indiscretion was to prove hugely pivotal. Just two minutes later, Italy closed to within a point when debutant Louis Lynagh – son of former Australia international Michael – ran on to Garbisi’s kick-through and darted over the line. Garbisi – with the chance to edge his team in front – saw his conversion attempt come back off the post.

The Scots found themselves in trouble in the 57th minute when the Italians got themselves in front as replacement scrum-half Stephen Varney found a gap between Andy Christie and Jack Dempsey to nudge his way over. This time Garbisi was on target.

Garbisi then scored a huge penalty from distance to put the hosts nine points ahead with seven minutes to play, placing the Scots in a state of desperation.

Replacement lock Sam Skinner – denied a match-winning try against France a month previously – pushed over with two minutes left and Russell converted to bring the Scots back to within two points and set up a tense finish.

Townsend’s team were unable to find another score, however, and – as the Azzurri celebrated a rare and deserved victory – the Scotland players slumped to their knees in abject despair as a campaign had that promised so much was reduced to rubble in Rome.

Rory Darge feels Italy’s Stadio Olimpico would be “a nice place” for Duhan van der Merwe to become Scotland’s record try-scorer – but the co-captain stressed that his team’s focus is solely on leaving Rome with another Guinness Six Nations victory.

The Scots have won two of their three championship matches so far to sit second in the table – albeit six points adrift of Grand Slam-chasing Ireland – going into the penultimate round of fixtures.

While the priority for Gregor Townsend’s side is to get the points that will keep them on course for a first top-two finish in the Six Nations era, the fixture has added spice because Van der Merwe goes into it with 26 international tries to his name – one shy of the county’s record-holder Stuart Hogg.

A score or two by the South Africa-born wing on Saturday would see him join or overtake the former full-back, who will be at the Stadio Olimpico to support his old colleagues after completing a charity cycle to the Eternal City in aid of the My Name’5 Doddie foundation.

Back-rower Darge said: “I know one of Duhi’s tries in particular (his second one against England) was off our defensive efforts and then the turnover attack, so if we do get ourselves right, then we can put guys like that in.

“He’s obviously a great finisher. Sometimes you only need to give him half a chance and he comes away with a chance, so it (the record) could happen.

“It would be a nice place for him to do it but I don’t think he’ll have too much of his mind set on that.

“He’ll just be focused on his role, and he might come away with a couple, but we’ll have to wait and see. I’m sure Italy will put us under a lot of pressure.”

Darge insisted Edinburgh wing Van der Merwe has remained as humble as ever in the wake of the Calcutta Cup hat-trick a fortnight ago that put him on the cusp of Hogg’s record.

“Duhi’s always the same,” said the Glasgow forward. “Around training and around camp, he’s always quite laidback but when he gets out there he’s a Test-match animal, and it’s class to be able to play with him.”

Darge’s only previous experience of playing at Stadio Olimpico was a 33-22 victory two years ago in front of a crowd of 41,214.

This time, the famous arena in north-west Rome – used predominantly for football – is set to be full to capacity, with 70,000 tickets having been sold on the back of encouraging championship performances from the Azzurri against England and France.

“Italy are a really good side,” said Darge. “We saw that against France, and they’re obviously playing at home where they’ll have a really good atmosphere and good backing.

“The Stadio Olimpico is different (to other Six Nations venues) but the Scotland fans always travel well for this one, so that will be massive for us.

“It’s a long walk from the changing room but it’s a cool stadium. Inside the changing room, on the walk in, with all the jerseys on the wall and stuff like that, there’s a lot of history behind it, so it’s a cool place to be playing rugby.”

Scotland face Italy in their penultimate Guinness Six Nations match at the Stadio Olimpico on Saturday.

Here, the PA news agency looks at some key talking points ahead of the showdown in Rome.

History beckons for Duhan

There is every chance of Scottish history being made in Rome this weekend. Duhan Van Der Merwe moved within one of Scotland’s record try-scorer Stuart Hogg when he scored a match-defining hat-trick against England at Murrayfield last time out. A single score will be enough to take the South Africa-born wing level with Hogg on 27, but in his current form – five tries in his last three Six Nations matches – few would bet against him doing enough at the Stadio Olimpico to break the record outright. Hogg, incidentally, is set to be watching on from the stand after completing a fundraising cycle to the Italian capital in aid of the My Name’5 Doddie charity.

Scots in hunt for rare top-two finish

Scotland have not finished in the top two of the championship since winning the last staging of the Five Nations in 1999. They arrive in Rome knowing a victory will keep them on course to do so for the first time in the 21st century. Gregor Townsend’s team are currently second in the table – a point ahead of England and three above France – with two matches to play. While they still have an outside chance of pipping Ireland to the title, finishing second appears to be a more realistic target.

Chance for Scots trio to shine

There is an element of freshness to the Scotland team this weekend as Townsend has handed opportunities to three players who have been regular squad members in recent seasons but who are not accustomed to starting in the dark blue. Burgeoning Bath centre Cam Redpath makes his fourth Scotland start in place of the injured Sione Tuipulotu, Glasgow scrum-half George Horne has been handed his first start since the 2019 World Cup as Toulon’s Ben White gets the weekend off to rest, while in-form Saracens back-rower Andy Christie has finally been rewarded with first international start two years after making his debut at home to France.

Italy a sticky opponent

Italy have bounced back well from a chastening World Cup and have proved particularly testing opponents for England and France under recently-installed head coach Gonzalo Quesada. The Azzurri lost by just three points to the Red Rose on match-day one and drew away to France in their last outing, with a last-gasp penalty miss costing them what would have been a famous victory in Lille. The Italians have lost their last 13 meetings with the Scots but they gave Townsend’s team a good game at Murrayfield a year ago and have every reason to be confident of doing so again on home soil this weekend.

Full house at the Olimpico

The Stadio Olimpico regularly hosts capacity crowds for football matches, such as Thursday’s Europa League showdown between Roma and Brighton, but it is far more unusual for the iconic bowl in the north-west of the Eternal City to sell out for rugby matches. It is a sign of the relative buoyancy of both of these nations, therefore, that almost 70,000 tickets have been sold for Saturday’s match.

Gregor Townsend is desperate to see Scotland hit their maximum performance levels over the next two weekends as they bid to secure a first top-two finish in the Six Nations era.

The Scots are second in the championship table – a point ahead of England and three above France – as they prepare for matches away to Italy this Saturday and Grand Slam-chasing Ireland the following week.

Townsend has seen his team defeat Wales and England while going agonisingly close to defeating the French, but he still feels they are yet to hit top gear for any sustained period.

“Happy-ish,” he said, reflecting on the campaign so far. “You’ve obviously got to be delighted for the team to beat England and retain the Calcutta Cup.

“But in terms of performance, we’re a little bit away from what we know we can deliver. The first 45 minutes against Wales is probably still the best we’ve played.

“We had elements of control throughout much of the France game and for the last 60 minutes of the England game. But there’s still a lot more to come from us.

“We’re delighted with how the guys trained yesterday, a real physical session. The non-23 trained really well and really tested us. So the guys are in a good position to deliver their best performances over the next two weeks. That’s all we’re working towards.”

Scotland know that even if they win in Rome on Saturday, it may not be enough to stop Ireland securing the championship against England later in the day. If the Irish slip up at Twickenham, it could pave the way for a last weekend title shootout between Ireland and Scotland in Dublin.

“We can only do what we have to do this weekend which is deliver a winning performance and see where we are after that,” said Townsend.

“We’re the first game so I’m sure we’ll be watching the other game with interest.

“We’re into the last two rounds of the championship still in the mix, although it’s an outside chance for the championship. There’s a lot to play for this weekend and again in Dublin.”

Asked if a second-place finish would represent a good outcome for the Scots, who have never finished higher than third this century, Townsend, said: “I don’t know. Anything to build on what we’ve done in the last few years would be an improvement.

“We’re focused on the performance and we know that at times it’s been better since the World Cup and last year’s Six Nations but still not where we believe this team can be.”

Townsend has made three changes to his team for the match in the Stadio Olimpico.

Regular scrum-half Ben White has been rested following a busy schedule with Toulon, paving the way for Glasgow’s George Horne to make his first start since the 2019 World Cup and the experienced Ali Price – who has played no part in the championship so far – is promoted to the bench.

In-form Saracens back-rower Andy Christie replaces Jamie Ritchie, who drops to the bench, while Bath’s Cam Redpath takes the place of influential Glasgow centre Sione Tuipulotu, who is out for the remainder of the tournament with a knee injury.

“We feel Ben has played a lot of rugby, he’s only missed one game since October,” said Townsend, explaining the changes at scrum-half.

“Going back and forward to Toulon during this period, something’s got to give in terms of a rest at some point, and we feel this week is the best week for him to recharge.

“And also with the form and experience of the two other scrum-halves in our squad who have been training really well. George has been on the bench and Ali has been excellent during this period so we want to give those two an opportunity.”

Scrum-half George Horne has been handed a surprise start for Scotland’s Guinness Six Nations showdown with Italy in Rome on Saturday, with Gregor Townsend making three changes to his starting XV.

The 28-year-old Glasgow back has been given the nod to wear the number nine jersey for the first time since the 2019 World Cup. Regular starter Ben White is rested and drops out of the squad altogether, with Ali Price – who has played no part in the tournament – promoted to the bench.

Saracens flanker Andy Christie is handed his first international start in place of Jamie Ritchie, who is among the substitutes.

Bath centre Cam Redpath made a positive impact off the bench in the 30-21 victory over England and replaces Glasgow’s Sione Tuipulotu, who will miss the remainder of the championship with a knee injury.

Back-rower Matt Fagerson, who dropped out of the 23 altogether for the last match, returns to the bench, with Townsend switching from a 5/3 split to a 6/2.

The Scots are currently second in the table and need a win to keep alive their slim championship hopes.

Craig Gordon is adamant he would feel equipped to go to this summer’s European Championships with Scotland even if he is unable to win back his Hearts starting place from Zander Clark.

The 41-year-old was undisputed number one for club and country before suffering a double leg-break on Christmas Eve of 2022 that sidelined him for almost a year.

Gordon returned to contention in December but has so far been restricted to just two Scottish Cup appearances against lower-league sides Spartans and Airdrie, with Hearts boss Steven Naismith keeping faith with Clark for cinch Premiership duties.

With the Euros kicking off in just over three months, the former Sunderland and Celtic keeper is running out of time to re-establish himself at club level.

Asked if he feels he would be fit and sharp enough to go to the showpiece in Germany even if not playing regularly for Hearts, Gordon – capped 74 times by his country – said: “Absolutely, yes. I feel really good.

“My body’s in the best shape it has been in years. There’s no doubt in my mind I’m capable of doing that. I just hope I get the opportunity to show everybody else as well.

“I’ve not spoken to Steve Clarke but I’m still hopeful. I still feel as if I’m capable, I feel my body’s in a good place, I feel ready to play. I definitely feel like I can still play at that level but that’s a choice for Steve.”

Gordon is expected to get another timely run-out for Hearts in Monday’s Scottish Cup quarter-final away to Morton, with the Scotland squad for the March friendly double-header against Netherlands and Northern Ireland due to be named just days later.

The veteran appears to be vying with current number one Angus Gunn, clubmate Clark, Motherwell’s Liam Kelly and Rangers number two Robby McCrorie to make the squad.

“We’ll see,” said Gordon, regarding the prospect of being selected by Clarke next week. “I’ll hopefully get the nod for the game on Monday (at Morton), and if I continue playing in the cup, it gives me another appearance, another chance to play.

“I don’t know what the manager (Clarke) is thinking but I’m certainly ready and able to join up if he decides to choose me.”

Asked if he could envisage a scenario whereby both he and clubmate Clark go to the Euros, Gordon said: “Anything’s possible. I don’t know what the manager’s thinking, what he sees as his best options but I’m still hopeful.

“There are nine league games to go, potentially three cup games, so I’ve got a lot still to fight for. I need to play as many of those as possible and try and force my way in there.”

Gordon has generally been a number one throughout his career and admits he is in a “strange” situation whereby he finds himself cast as a back-up despite never having been dropped due to his form at Hearts.

“Being a goalkeeper, it seems to change things slightly in terms of people’s views,” he said. “There’s two sides to it. If you’re in my camp, you say I only lost my place because of an injury, and if you’re not then you say the man in possession keeps the gloves.

“There’s two ways to think about it. It’s not fallen on my side just yet but I can only do what I can do. I feel like I’m training really well and giving myself that opportunity.”

*Craig Gordon was speaking as Hearts launched their Maroon Mile project to mark 150 years of history in Edinburgh’s Gorgie and Dalry area. The Jambos have announced plans for a trail dedicated to their existence and achievements within the local community.

Doug Watson has been appointed as Scotland’s new head coach, returning to the role he performed on an interim basis last year.

The South African stepped in when Shane Burger departed for Somerset and successfully steered the side to victory in the T20 World Cup qualifier.

He will now be the man to take charge of that tournament in the West Indies this summer, where Scotland will take on England, Australia, Oman and Namibia in the group stages.

The 50-year-old takes up his post in April after completing the season with Auckland, who he has already led to the 20-over Super Smash title.

“I’m really excited to be rejoining Cricket Scotland again. I had a wonderful time here in 2023 and I’m looking forward to getting back to work with the players and staff alike,” he said.

“I love the city of Edinburgh and the Scottish people, and working with such a good group of players and seeing their determination and their passion to represent Scotland and the badge was a big part of my desire to come back.

“We’ve got a lot of hard work to do ahead of the T20 World Cup, but there’s some great fixtures to look forward to. We play England and Australia, as well as Namibia and Oman, so they’re all going to be tough games, but we’ve also got to make sure that we see it as a wonderful opportunity to showcase our passion, our fight, and our skills.”

Steve Snell, Cricket Scotland’s head of performance said: “We interviewed candidates across the world from a number of high-performance environments and it was a hugely competitive process.

“Throughout, it was clearly evident that Doug is a high-quality coach with experience across a variety of environments. He is an excellent cultural fit for Cricket Scotland. He will make a huge difference to the game here, not just to the Scotland men’s squad but also within Scottish cricket as a whole.”

Joe Marler is desperate to help England wrestle back the Calcutta Cup on Saturday after growing exasperated with Scotland’s recent dominance of the fixture.

The 33-year-old prop grew up in an era when the Red Rose firmly held the upper hand over the Scots and he was on the winning side four times in a row after first playing in the highly-charged showdown in 2014.

The tables have turned since 2018, however. Scotland have lost only one of their last six matches against the Auld Enemy under Gregor Townsend and go into this weekend’s match buoyed by having won each of the last three.

That situation rankles with Marler, who is intent on ensuring England are celebrating on enemy territory come Saturday evening.

“It would just be nice to be on the winning end of it for once because it has been so long since we have,” he said at Murrayfield on the eve of the match.

“Obviously we had 2020, but the continued dominance from Scotland over us – it has been a long time now.

“From the start, we hadn’t lost to Scotland. Then the 2018 game the tide started to turn, the players that have come through in the Scottish side, you go, ‘Hang on, they have got some world-class operators now’.

“And it does shift the mindset slightly coming in as underdogs.

“The last few years without that cup, seeing Finn Russell and Greig Laidlaw, that video of them with their shirts off and singing with the cup.

“I wish I could be doing that rather than watching it. Or that famous Finn Russell photo where he’s got his Spiderman hands up and he’s loving it.

“That stirs passion in me to go, ‘I want that cup’, and I know a number of the other boys in the team want that cup back as well.”

Marler is pleased to still be in a position to help improve England’s record against Scotland after wondering if the World Cup last autumn might signal the end of his international career before Steve Borthwick assured him he still had a part to play.

“It’s almost like an addiction,” he said of his ongoing involvement with the national team. “I want to be part of a winning England team, creating new stories, creating new memories.

“I thought the World Cup was going to be my last opportunity at that, but Steve rang to ask if I’m still hungry.

“He asked if I had the desire to still crack on because he needed to blood some new players and move into the next cycle, but he also needed guys around to help with that.

“I asked my wife first but for me, yeah, it’s that addiction to be part of a winning England team and helping those young guys come through. That’s why I keep coming back.”

England will be led out at Murrayfield on Saturday by captain Jamie George, who lost his mother, Jane, a week last Wednesday following a short battle with lung cancer.

Marler, who will start on the bench, has been hugely impressed with the way his friend and fellow 33-year-old front-rower has dealt with the situation.

“Jamie has been incredible,” said the Harlequins prop. “I remember talking to him a few weeks before coming into camp, and he was talking about the captaincy being offered to him and he wasn’t sure whether he was going to take it up with things going on with his mum.

“Having known Jane since I was 16, 17, coming through the age groups with Jamie, I said, ‘Mate, just flip it and tell her you’re not doing it and see how upset, disappointed and gutted she’d be if you didn’t do it. You’ve got enough support around you in terms of the senior group to help you with it and you’re the best bloke for it, so let’s crack on and do it.’

“I’m pleased that he did. It has been tough since we found out about Jane, but he’s shown huge strength, and vulnerability which is great for the whole group, especially the youngsters to see that.

“He’s very much thinking, ‘I want to come up here, get the job done’. He’s dealt with it incredibly well.”

Scotland talisman Finn Russell is unfazed by the prospect of being targeted by England’s new blitz defence in Saturday’s Calcutta Cup showdown at Murrayfield.

The Red Rose have adopted a more aggressive approach for this year’s Guinness Six Nations after highly-regarded defence coach Felix Jones joined Steve Borthwick’s backroom team in the wake of helping South Africa win the World Cup. 

England are expected to try to swarm stand-off Russell and his midfield colleagues in an effort to neutralise Scotland, but the 31-year-old has no issue with the possibility of being singled out.

“It’s probably similar to a lot of teams in that the 10s are the key players in attack,” said co-captain Russell. “I’m not sure what England are going to do – if they are going to fire out the line and try to take me out or shut me down from the outside.

“That is something we will have to figure out in the game. We will have to be able to adapt, with myself and Sione (Tuipulotu) and Huw (Jones) being on the same page and having Blair (Kinghorn) out wide as another option.

“Although the 10 controls a lot of the attack, it is not just down to me to create things. We will be looking to other boys to get away from them.”

Scotland were tamed the last time they came up against a Jones-inspired blitz defence when they lost 18-3 to South Africa at the World Cup in September, but Russell insists they have learned from that encounter.

“We have looked back and talked about that game, and obviously looked at England’s first two games of this campaign,” said Russell. “I think our learnings from the World Cup were not to go into our shells if we do feel the pressure.

“There were chances in that game against South Africa that we probably never saw on the pitch. Under pressure we probably went into our shell a little bit.

“Tomorrow we just need to have belief in ourselves and trust the work we have put in over the last six months to a year.

“At times we will be under pressure and it will be tough, but we can fall back to what we have done building up to this game.

“We can have belief and confidence in ourselves and hopefully we can take the chances that will be out there.”

After Russell lost his first three Calcutta Cup matches, including a 61-21 defeat at Twickenham in 2017, the Scots have won each of the last three meetings and have lost only one of the last six.

“With us and England, we have been progressing and over the last few years they have potentially not been as good as they can be,” said Russell. “But the World Cup showed how good they can be, getting to the semi-finals.

“Obviously they have won their first two games of this campaign so they are getting back to where they should be. They are one of the best teams in the world.

“We can’t look back at the last few games and think it has turned in our favour. Every time we play England, it is always a huge challenge and we have got to be at our best to be able to beat them.”

Scotland and England renew hostilities when they meet in rugby’s oldest international fixture at Murrayfield on Saturday.

For both sides it is win or bust in a critical Guinness Six Nations round-three clash that will shape their respective Championships.

Here, the PA news agency looks at five talking points heading into the Edinburgh showdown.

Furbank’s second coming

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Steve Borthwick is capable of throwing curve balls in selection – think Marcus Smith at full-back, Alex Mitchell starting at scrum-half at the World Cup – and the latest example is George Furbank’s return at full-back. Freddie Steward controls the air against any opposition but Borthwick has cast aside England’s safety blanket in favour of a more natural ball player who will provide a counter-attacking threat and greater mobility in defence. Promoting Furbank is a bold call and even if the six caps won between 2020 to 2022 failed to provide compelling evidence of his Test pedigree, he is an exciting pick who has been on fire for Northampton this season.

Heavyweights collide

England take a more balanced backline to Edinburgh after recalling Ollie Lawrence at inside centre. For the first time in this Six Nations there will be genuine ball-carrying clout in midfield after Lawrence recovered from the hip injury that ruled him out of the wins against Italy and Wales. In the words of assistant coach Kevin Sinfield, the powerful Bath runner is “ready to kick the door down” and it will be hoped he can provide a counter weight to the similarly physical Sione Tuipulotu. It is a heavyweight collision that will influence the outcome of the match.

Lopsided rivalry

Scotland are odds-on favourites to retain the Calcutta Cup – and rightly so. They have won their last three Tests against the ‘Auld Enemy’, whose victory in 2020 is their only triumph in the last six meetings. The games have been ferociously competitive but Scotland are simply a better team, having turned a one-sided rivalry on its head. Defined by this fixture, these are the glory days for Scottish rugby and for a measure of England’s decline in recent years – finishing third at the 2023 World Cup aside – look no further than their recent struggles against their oldest foes.

Moment of truth

Gregor Townsend admitted that the history and emotion of a clash with England made it Scotland’s “most important game of the season”, but the head coach also knows that settling old scores is only part of the bigger picture. A golden generation in the nation’s rugby history, epitomised by their fly-half genius Finn Russell, is in danger of passing without winning any silverware and after the injustice of seeing a late match-winning try disallowed against France in round two, they can not afford any more slip ups. Time is running out for Russell’s Scotland to prove they are a serious team.

Cautious optimism

England arrive at Murrayfield with two wins in the bank and alongside Ireland they are the only unbeaten team left in the tournament. Coupled with their bronze medal finish at the World Cup and that should be cause for optimism when they face Scotland for the 142nd time. But a side in transition that is attempting to evolve its attack and get to grips with a new blitz defence has so far faced the Six Nations’ two weakest sides. The level of competition cranks up significantly on Saturday and while there is no danger of Borthwick’s resilient side being blown away, defeat would signpost another Championship of underachievement.

Jamie George will draw inspiration from the heartwarming thought that his late mother will be watching down on him as he prepares to lead England into Saturday’s Calcutta Cup showdown with Scotland just over a week after her death.

The 33-year-old Saracens hooker found out on the same day that he was appointed captain of the Red Rose last month that his mum Jane had been diagnosed with cancer. Her situation deteriorated quickly and she died last Wednesday.

George takes some solace from the fact a woman he described as “the biggest rugby fan on earth” was able to see her boy skipper his country for two matches, the Guinness Six Nations victories over Wales and Italy.

“We’ve been going through a lot as a family for a long period,” he said, speaking with remarkable composure about his ordeal from England’s team hotel in Edinburgh city centre on Thursday evening.

“The deterioration she had was really fast. I found out on Sunday about the fact that she was terminal, and she passed away on Wednesday [last week].

“My mum was the biggest rugby fan on earth, she loved this team, loved watching me play, she never missed a game.

“The text I’ve got from her before my first game as captain is something I will treasure forever. She said it was the proudest day of her life so given what she was going through, to still be able to put a smile on her face was huge.”

George was adamant he did not want to excuse himself from England duty. He turned up on the Friday after his mum’s death to participate in an open training session at Twickenham and had no doubt in his mind that he wanted to lead his team into battle with Scotland.

George’s father, his brothers, his uncle and his cousin will be at Murrayfield for what he hopes will be a cathartic experience for the family amid the trauma.

“Taking time off is the last thing she would have wanted me to do,” said George. “It’s not what I wanted to do.

“I feel very privileged to do what I do and hopefully the boys will agree that I’ve been able to fulfil my role as captain and fulfil my role as a player in this team.

“It’s not an ideal situation to be in, but there was never any doubt in my mind that I wanted to be involved in this game.

“Wherever she is now, she will be looking down telling everyone that is there that her son is the England captain. I know for a fact that meant a huge amount to her.

“Whenever I’ve played, I’ve always wanted to make my family proud. It’s been a huge driver for me. That won’t change this weekend – it will probably be enhanced this weekend.

“It will be emotional for me coming out. It will be the first game that she won’t be there. She wasn’t able to come to the first two games to watch, which has been tough in itself, but before that she was always there, she never missed it.

“My dad, my uncle, my cousin and both brothers are coming up this weekend. It’s going to be great for them to be able to be there. It’s amazing what rugby can do in situations like this.

“When I first became captain, I spoke a lot about showing how much it means to play for England and what an amazing impact you can have on people’s lives.

“I have seen it first-hand because my mum was on her death bed talking about the England rugby team and how proud she was of me being able to do what I do.

“That’s absolutely incredible. She will be with me in some capacity on Saturday and that means a huge amount to me.”

Gregor Townsend says Scotland’s sole focus is on winning their “biggest game of the season” against England this weekend and maintaining their recent dominance of the Calcutta Cup.

The Scots go into Saturday’s showdown with their bitter rivals knowing they will almost certainly require a victory to stay in contention for the Guinness Six Nations title after their controversial defeat by France last time out.

Townsend is adamant that any lingering injustice from ‘trygate’ a fortnight ago has been parked and that any talk of contending for the title can remain firmly on the backburner until after their high-stakes encounter with Steve Borthwick’s side.

“When you’re coming in off the back of a defeat, you have to bounce back with a victory,” said Townsend, when asked on Thursday how the France defeat has left his side’s title prospects.

“You’re not really thinking about the championship. We’ll see where we are on Saturday.

“The full focus is on this game because it’s our most important game of the season.

“It’s the most important game for our supporters and we also play for a trophy, in amongst the Triple Crown and other trophies.

“But this is one that we’re focused on more than the others. That probably shouldn’t be the case, but it is.

“It’s history. It’s emotion. We’ve seen the impact this game has on Scottish supporters when we do manage to get a victory in this fixture. That’ll drive us on Saturday.”

Having beaten England only three times in 27 attempts between 1990 and 2018, the Scots now find themselves going into this weekend’s fixture on the back of a three-game winning run and having lost only one of their last six meetings with their old foes.

Townsend, who was accustomed to regular defeats against England in his time as a player, admits his team’s recent burst of Calcutta Cup success has given them increased belief going into Saturday’s match.

“Yes, for sure,” he said. “The game in 2018 (Scotland’s first win over England in a decade) has certainly given the players confidence when they’ve taken on England in the last few games.

“But what’s most relevant is the game you’ve just played, the things you have to work on to be a better team, and the threats that England bring.

“This England team is different to the one we faced 12 months ago but ultimately it’s just about delivering in the 80 minutes on the day.”

Townsend has made three changes to the side that started the 20-16 defeat by France, with Glasgow wing Kyle Steyn returning after missing Les Bleus game when his wife went into labour and Toulouse full-back Blair Kinghorn back after sitting out the first two matches with a knee injury.

Kyle Rowe and Harry Paterson, who deputised in the absence of the two experienced backs, drop out of the squad altogether.

The most notable change is in the back row where Edinburgh flanker Jamie Ritchie, who recently lost the captaincy and then was then left out of the 23 for the France game, returns to the number six jersey in place of Matt Fagerson, who is dropped from the squad.

“Jamie has really accepted the challenge that was there about a month ago that there’s increased competition in the back-row and for that reason he was no longer going to be captain,” explained Townsend.

“But since that conversation he’s played two games for Edinburgh and one for Scotland in Cardiff and he’s played well.

“We believe this game will suit his strengths and his experience he can bring to the team as well will be a boost.”